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Bedeutung von Isolation, Flaechengroesse und Biotopqualitaet. Modellbildung und Simulation von Austauschprozessen, Mikroevolution und Extinktionsrisiko

CODENET: Coccolithophorid Evolutionary Biodiversity and Ecology Network

1. Evolutionary biodiversity: A study of diversity of coccolithophorid biology relative to higher level phylogeny. Objectives: - Determine the major patterns of biodiversity in coccolithophorid life-cycles, biomarker composition, photosynthetic pigments, cytology and plastid genome.- Re-evaluate the phylogeny of the coccolithophorids using separate and combined analyses of: molecular genetic, morphological, biochemical and palaeontological data. - Calculate divergence times of groups and rates of evolution, including molecular clock calibrations. - Reconstruct the sequence of major evolutionary steps in coccolithogenesis, lipid biochemistry, plastid evolution, and life cycle differentiation. 2. Microevolution and species-level variation: A study of microevolutionary pattern and process, species concepts, and the significance of fine-scale genotypic diversity. Objectives: - Evaluate which aspects of variation represent genotypic vs. ecophenotipic or ontogenetic variation. - Determine whether intra-specific variability in morphology, physiology and biochemistry are correlated, defining discrete sub-species. - Determine whether microevolution occurs by (sub-)species selection or effectively sympatric evolution within ocean-scale populations. 3. Ecology: An investigation of the prime constraints on coccolithophorid ecology and the ecological significance of biodiversity within the coccolithophorids. Objectives: - Determine whether coccolithophorids as a group occupy a distinctive ecological niche, and if so characterise it. - Determine which aspects of intra- or inter-specific assemblage variation are most valuable for palaeoecological calibrations and develop palaeoecological proxies. - Determine the extent to which coccolithophorid carbonate accumulation rate is affected by species composition and evolution.

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