*Der Gesundheitszustand der Bäume im Schweizer Wald wird seit 1985 mit der Sanasilva-Inventur repräsentativ erfasst. Die wichtigsten Merkmale sind die Kronenverlichtung und die Sterberate. Das systematische Probeflächen-Netz der Inventur ist im Laufe der Zeit ausgedünnt worden. In der Periode von 1985 bis 1992 wurden rund 8000 Bäume auf 700 Flächen im 4x4 km-Netz aufgenommen, 1993, 1994 und 1997 rund 4000 Bäume im 8x8 km-Netz und in den Jahren 1995, 1996 und 1998 bis 2002 rund 1100 Bäume im 16x16 km-Netz . Aufnahmemethode Alle drei Jahre (1997, 2000) wird die Sanasilva-Inventur auf dem 8x8-km Netz (ca. 170 Probeflächen ) durchgeführt. In den Jahren dazwischen findet die Inventur auf einem reduzierten 16x16-km Netz (49 Probeflächen) statt. Jede Fläche besteht aus zwei konzentrischen Kreisen. Der äussere Kreis hat ein Radius von 12.62 m (500 m2) und der innere ein Radius von 7.98 m (200 m2). Auf dem inneren Kreis werden alle Bäume mit einem Mindestdurchmesser in Brusthöhe von 12 cm und auf dem äusseren Kreis mit einem Mindestdurchmesser in Brusthöhe von 36 cm aufgenommen. In Nordrichtung wird zusätzlich in 30 m Entfernung eine identische Satellitenprobenfläche eingerichtet. Die Aufnahme findet in Juli und August statt. Eine Aufnahmegruppe besteht aus zwei Personen, von denen eine die Daten erhebt, und die andere die Daten eintippt. Die Daten werden mit dem Feldkomputer Paravant und der Software Tally erfasst. Die Aufgabenteilung wechselt zwischen Probeflächen. Auf dem 8x8-km Netz werden zusätzlich 10 Prozent der Flächen von einer unabhängigen zweiten Aufnahmegruppe zu Kontrollzwecken aufgenommen. Hauptmerkmale der Sanasilva-Inventur: Die Sanasilva-Inventur erfasst vor allem folgende Indikatoren des Baumzustandes: Die Kronenverlichtung wird beschrieben durch den Prozentanteil der Verlichtung einer Krone im Vergleich zu einem Baum gleichen Alters mit maximaler Belaubung/Benadelung an diesem Standort, den Anteil dieser Verlichtung, der nicht durch bekannte Ursachen erklärt werden kann, den Ort der Verlichtung, den Anteil und den Ort von unbelaubten/unbenadelten Ästen und Zweigen. Die Kronenverfärbung wird durch die Abweichung der mittleren Farbe (aufgenommen als Farbton, Reinheit und Helligkeit nach den Munsell Colour Charts) eines Baumes zu der für diese Baumart typischen Normalfarbe (Referenzfarbe) und durch das Vorhandensein, das Ausmass und den Ort der von der Referenzfarbe abweichenden Farben beschrieben. Der Zuwachs eines Baumes wird durch die zeitliche Veränderung der aufgenommen Baumgrössen beschrieben (Brusthöhendurchmesser, Höhe des Baumes, Kronenlänge und Kronenbreite). Weitere Merkmale sind die erkannten Ursachen der Kronenverlichtung, die Kronenkonkurrenz und das Vorkommen von Epiphyten, Mistel und Ranken in der Baumkrone.
Introduction: In Malaysia, excessive nutrients from livestock waste management systems are currently released to the environment. Particularly, large amounts of manure from intensive pig production areas are being excreted daily and are not being fully utilised. Alternatively, the excess manure can be applied as an organic fertiliser source in neighbouring cropping systems on the small landholdings of the pig farms to improve soil fertility so that its nutrients will be available for crop uptake instead of being discharged into water streams. Thus, there is a need for better tools to analyse the present situation, to evaluate and monitor alternative livestock production systems and manure management scenarios, and to support farmers in the proper management of manure and fertiliser application. Such tools are essential to quantify, and assess nutrient fluxes, manure quality and content, manure storage and application rate to the land as well as its environmental effects. Several computer models of animal waste management systems to assist producers and authorities are now available. However, it is felt that more development is needed to adopt such models to the humid tropics and conditions of Malaysia and other developing countries in the region. Objectives: The aim is to develop a novel model to evaluate nutrient emission scenarios and the impact of livestock waste at the landscape or regional level in humid tropics. The study will link and improve existing models to evaluate emission of N to the atmosphere, and leaching of nutrients to groundwater and surface water. The simulation outputs of the models will be integrated with a GIS spatial analysis to model the distribution of nutrient emission, leaching and appropriate manure application on neighbouring crop lands and as an information and decision support tool for the relevant users.
Magnetic resonance tomography (MRT) on microcosm soil cores (200 mm Ø) used for CeMiX, comprising naturally stacked subsoil down to 700 mm plus topsoil from CeFiT, will be implemented at a laterally partially open Split 1.5 T magnet, with intended final in-plane spatial resolution of 200 Micro m. Three-dimensional biopore distributions and dynamics of their formation within the cores will be determined non-invasively and compared to complementing CT analyses of SP 2. One major aim is a non-invasive differentiation of the biopores into earthworm- and root system-originating ones and currently air-, water-, root- and earthwormfilled ones, based on NMR relaxation parameters. Attempts will additionally be made to classify different wall coatings of the biopores with regard to their water affinity. Dynamics of water distribution within the microcosm core and its biopore structures, starting from initial values taken from CeFiT (SP 3), will be documented with an in-plane resolution of 5 mm, in parallel to measurements of root growth dynamics for calculation of biomass and root surface area. Special emphasis will be put on the role of the plant root system for a re-distribution of water/D2O (and solutes) between different soil layers. Finally we will attempt MRT-controlled sample collection from the microcosm cores, to get - together with our research unit partners of SPs 4-8 - repeated access to minimally invasively acquired data on nutrient and microorganism distributions in concert with non-invasively collected water and root distribution data as a basis for dynamic modelling of water and solute circuits in SP 10. Beside the microcosm cores, flat rhizotrons as used in SP 3 will be employed to enable measurements of root and shoot hydrostatic pressure profiles with pressure probes, in addition to MRT measurements. In this way water distributions and corresponding driving forces and growth dynamics will be measured altogether in a minimally invasive manner.
It has been suggested that dying and decaying fine roots and root exudation represent important, if not the most important, sources of soil organic carbon (SOC) in forest soils. This may be especially true for deep-reaching roots in the subsoil, but precise data to prove this assumption are lacking. This subproject (1) examines the distribution and abundance of fine roots (greater than 2 mm diameter) and coarse roots (greater than 2 mm) in the subsoil to 240 cm depth of the three subsoil observatories in a mature European beech (Fagus sylvatica) stand, (2) quantifies the turnover of beech fine roots by direct observation (mini-rhizotron approach), (3) measures the decomposition of dead fine root mass in different soil depths, and (4) quantifies root exudation and the N-uptake potential with novel techniques under in situ conditions with the aim (i) to quantify the C flux to the SOC pool upon root death in the subsoil, (ii) to obtain a quantitative estimate of root exudation in the subsoil, and (iii) to assess the uptake activity of fine roots in the subsoil as compared to roots in the topsoil. Key methods applied are (a) the microscopic distinction between live and dead fine root mass, (b) the estimation of fine and coarse root age by the 14C bomb approach and annual ring counting in roots, (c) the direct observation of the formation and disappearance of fine roots in rhizotron tubes by sequential root imaging (CI-600 system, CID) and the calculation of root turnover, (d) the measurement of root litter decomposition using litter bags under field and controlled laboratory conditions, (e) the estimation of root N-uptake capacity by exposing intact fine roots to 15NH4+ and 15NO3- solutions, and (f) the measurement of root exudation by exposing intact fine root branches to trap solutions in cuvettes in the field and analysing for carbohydrates and amino acids by HPLC and Py-FIMS (cooperation with Prof. A. Fischer, University of Trier). The obtained data will be analysed for differences in root abundance and activity between subsoil (100-200 cm) and topsoil (0-20 cm) and will be related to soil chemical and soil biological data collected by the partner projects that may control root turnover and exudation in the subsoil. In a supplementary study, fine root biomass distribution and root turnover will also be studied at the four additional beech sites for examining root-borne C fluxes in the subsoil of beech forests under contrasting soil conditions of different geological substrates (Triassic limestone and sandstone, Quaternary sand and loess deposits).
In many plant species, FLOWERING LOCUS T and related proteins are the mobile signal that communicates information on photoperiod from the leaves to the shoots, where the transition to flowering is realized. FT expression is tightly controlled at the transcriptional level so that it is restricted to leaves, occurs only in appropriate photoperiods, and integrates ambient temperature and developmental cues, as well as information on biotic and abiotic stress. We previously established that FT transcription in the model plant Arabidopsis thaliana requires proximal promoter cis-elements and a distal enhancer, both evolutionary conserved among Brassicacea species. In addition, FT transcription is blocked prior vernalization in biannual accessions and vernalization-dependency of FT is controlled through a CArG-box located in the first intron that binds the transcriptional repressor FLOWERING LOCUS C (FLC). Chromatin-mediated repression by the Polycomb Group (PcG) pathway is required for photoperiod-dependent FT regulation and participates in FT expression level modulation in response to other cues.In this project, I propose to explore the available sequence data from the 1001 genome project in Arabidopsis to evaluate how often changes in regulatory cis-elements at FT have occurred and how these translate into an adaptive value. Allele-specific FT expression pattern will be measured in F1 hybrids of different accessions in response to varying environmental conditions. FT alleles that show cis-regulatory variation will be further analyzed to pinpoint the causal regulatory changes and study their effect in more detail. The allotetrapolyploid species Brassica napus is a hybrid of two Brassiceae species belonging to the A- and C-type genome, which are in turn mesopolyploid due to a genome triplication that occurred ca. 10x106 years ago. We will determine allele-specific expression of FT paralogs from both genomes of a collection of B. napus accessions. The plants will be grown in the field in changing environmental conditions to maximize the chance to detect expression variation of the paralogs. We will compare the contribution of the founder genomes to the regulation of flowering time and asses variation in this contribution. A particular focus will be to study the impact of chromatin-mediated repression on allele selection in B. napus.
Background and Objectives: The project area is located in the Ashanti Region of Ghana / West Africa in the transition zone of the moist semideciduous forest and tropical savannah zone. Main land use in this region is subsistence agriculture with large fallow areas. As an alternative land-use, forest plantations are under development by the Ghanaian wood processing company DuPaul Wood Treatment Ltd. Labourers from the surrounding villages are employed as permanent or casual plantation workers. Within three forest plantation projects of approximately 6,000 ha, DuPaul offers an area of 164 ha (referred to as Papasi Plantation) - which is mainly planted with Teak (Tectona grandis) - for research purposes. In return, the company expects consultations to improve the management for sustainable timber and pole production with exotic and native tree species. Results: In a first research approach, the Papasi Plantation was assessed in terms of vegetation classification, timber resources (in qualitative and quantitative terms) and soil and site conditions. A permanent sampling plot system was established to enable long-term monitoring of stand dynamics including observation of stand response to silvicultural treatments. Site conditions are ideally suited for Teak and some stands show exceptionally good growth performances. However, poor weed management and a lack of fire control and silvicultural management led to high mortality and poor growth performance of some stands, resulting in relative low overall growth averages. In a second step, a social baseline study was carried out in the surrounding villages and identified landowner conflicts between some villagers and DuPaul, which could be one reason for the fire damages. However, the study also revealed a general interest for collaboration in agroforestry on DuPaul land on both sides. Thirdly, a silvicultural management concept was elaborated and an improved integration of the rural population into DuPaul's forest plantation projects is already initiated. If landowner conflicts can be solved, the development of forest plantations can contribute significantly to the economic income of rural households while environmental benefits provide long-term opportunities for sustainable development of the region. Funding: GTZ supported PPP-Measure, Foundation
Die erste Antragsphase war auf die Bildungsraten und die Speicherung von anthropogenem Kohlenstoff (Cant) im Antarktischen Zwischenwasser (AAIW) fokussiert. Mit Hilfe von Freon (CFC) Daten konnten wir eine signifikante Reduktion der AAIW Bildungsrate von den 1990ern zu den 2000ern Jahren feststellen. Dies führte zu einer geringeren Steigerung der Cant Speicherung als vom atmosphärischen Cant Anstieg und einem unveränderten Ozean zu erwarten war. Um den Schwierigkeiten mit den Randbedingungen auszuweichen (Pazifisches AAIW strömt über die Drake Passage auch in den Atlantik und weiter in den Indischen Ozean) planen wir nun ein globales Vorgehen um in allen Ozeanen die Bildungsraten und Cant Speicherungen in den Zwischen- Tiefen- und Bodenwassermassen zu berechnen. Darüber hinaus wird der Zeitraum bis 2015 ausgedehnt, und wo immer die Datenlage es zulässt, Pentaden- anstatt Dekadenmittelwerte gebildet. Verwendet wird der aktualisierte GlODAPv2 Datensatz und eigene Daten.Die Berechnungen aus den Beobachtungen werden mit den Ergebnissen eines wirbelauflösenden globalen Ozeanmodells (1/10 Grad) kombiniert. Das POP Modell (Los Alamos Laboratory Parallel Ocean Program) mit eines horizontalen Auflösung von 0.1 Grad und 42 Tiefenstufen wird für die letzten 20 Jahre mit einem realistischen Forcing angetrieben und enthält außerdem die Freone als Tracer. Neben dem Vergleich mit einem klimatologischen Antrieb wird das Modell zur Weiterentwicklung der Tracer-Methode verwendet wir z.B. die Unsicherheit von zu wenig Datenpunkten und der Extrpolationsroutine auf die Bildungsraten / Cant Speicherungen. Ein weiterer wichtiger Punkt wird die Bestimmung der TTDs aus Lagrange Trajektorien und der Vergleich mit TTDs aus Tracermessungen sein, sowie die Untersuchung der Rolle der Wirbel, der Vermischung durch Wirbel und der vertikalen Vermischung.
Periods of extreme geomagnetic change such as geomagnetic excursions have frequently occurred throughout geological time. Characterizing their behaviour is essential for a full understanding of the geodynamo and the interaction of Earths magnetic field and the space environment. We propose to model the global behaviour of Earths magnetic field between 10 and 50 ka using palaeomagnetic data. During this time the geomagnetic field showed significant variability in direction and intensity, including two well documented excursions: Laschamp and Mono Lake. No model currently exists that spans the total length of this time period, yet this period could provide great insights into the geodynamo. The ultimate goal of the project is to synthesize the results from our empirical modelling with those from numerical dynamo simulations, so that a deeper physical understanding of geodynamo processes can be gained. We will compile all sedimentary and volcanic palaeomagnetic data coupled with geochronological data spanning this period. This data will be added to a community available database along with all rock magnetic and sedimentological metadata. This will allow a detailed assessment of the data used in the modelling. Low quality palaeomagnetic data and erroneous age models may distort geomagnetic field structures generated by our new model and it is a key objective of this study to assess the fidelity of the palaeomagnetic and chronological data included in the modeling. Using this data we will construct a temporally continuous global spherical harmonic geomagnetic field model through a regularized least squares inversion of the data using spherical harmonics in space and cubic B splines in time. This model will enable assessment of the geomagnetic at the core-mantle boundary, the Earths surface and at elevated altitudes. Our key scientific objective is to determine where excursions fit into the spectrum of geomagnetic field variations and how the geodynamo processes that generate excursions differ from those that produce secular variation and reversals.
(1) Terrestrische Biota der Antarktis sind durch geografische Isolation und inselhafte Verteilung geprägt. Die isolierte Lage der Antarktis und die Beschränkung auf weit voneinander entfernte kleine Habitatflecken haben zu einem hohen Endemiten-Anteil und einer starken Regionalisierung der Fauna und Flora geführt. Genetische Differenzierung, lokale Anpassung und die Evolution kryptischer Arten sind die Folge. Die Biodiversitäts-Konvention (CBD) betrachtet genetische Diversität als einen Eckpfeiler biologischer Vielfalt und stellt sie damit in eine Reihe mit der Diversität von Arten und Ökosystemen. Durch Einschleppung ortsfremder Arten und Homogenisierung bislang getrennter Genpools bedroht der Mensch jedoch zunehmend diese Isolation und genetische Differenzierung vieler antarktischer Biota. (2) Obwohl Flechten als wichtigste Primärproduzenten antarktische terrestrische Lebensräume dominieren, fehlen zurzeit Daten zu ihrer genetischen Struktur und Diversität. Der Umfang inter- und intrakontinentalen Genflusses ist bisher völlig unbekannt. Es ist deswegen derzeit unmöglich, den aktuellen und zukünftigen menschlichen Einfluss auf antarktische Flechtenpopulationen auch nur annähernd abzuschätzen.(3) Wir schlagen vor, mittels molekulargenetischer Daten die populationsgenetische Struktur von sechs weit verbreiteten Flechtenarten mit unterschiedlichen Ausbreitungsstrategien zu untersuchen. Dabei soll die Nullhypothese überprüft werden, dass Flechtenpopulationen genetisch nicht differenziert sind. Zusätzlich wollen wir abschätzen, ob menschliche Aktivitäten zur Einschleppung ortsfremder Arten oder Genotypen und zur Homogenisierung von Genpools beitragen. Hierfür sollen Lokalitäten mit hohem und niedrigem menschlichen Einfluss verglichen werden. Das Projekt schafft damit unverzichtbare Grunddaten für die Entwicklung von Schutzstrategien in der Antarktis.
Organic matter (OM) composition and dynamic in subsoils is thought to be significantly different from those in surface soils. This has been suggested by increasing apparent 14C ages of bulk soil OM with depth suggesting that the amount of fresh, more easily degradable components is declining. Compositional changes have been inferred from declining ä13C values and C/N ratios indicative for stronger OM transformation. Beside these bulk OM data more specific results on OM composition and preservation mechanisms are very limited but modelling studies and results from incubation experiments suggest the presence and mineralization of younger, 'reactive carbon pool in subsoils. Less refractory OM components may be protected against degradation by interaction with soil mineral particles and within aggregates as suggested by the very limited number of more specific OM analysis e.g., identification of organic compound in soil fractions. The objective of this project is to characterize the composition, transformation, stabilization and bioavailability of OM in subsurface horizons on the molecular level: 1) major sources and compositional changes with depth will be identified by analysis of different lipid compound classes in surface and subsoil horizons, 2) the origin and stabilization of 'reactive OM will be revealed by lipid distributions and 14C values of soil fractions and of selected plant-specific lipids, and 3) organic substrates metabolized by microbial communities in subsoils are identified by distributional and 14C analysis of microbial membrane lipids. Besides detailed analyses of three soil profiles at the subsoil observatory site (Grinderwald), information on regional variability will be gained from analyses of soil profiles at sites with different parent material.
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