Up to recent years, the study of trends and variability of the salinity (and freshwater) distributions was hampered by the lack of temporal and spatial resolution of the available observations (Wang DFG form 54.011 - 1/12 page 3 of 6 et al., 2010). With the onset of the Argo programme, the number of observed salinity profiles have significantly increased. The many Argo profiles distributed evenly over the year provide not only the means to calculate seasonally averaged salinity (and freshwater) distributions, but they can also be used to calculate regional Gravest Empirical Modes (GEMs) parameterized by pressure and dynamic height. Dynamic height is measured by altimetry and high quality altimeter data are available since 1992. GEMs will be used to construct from altimetry high resolution (in time and space) salinity, temperature and density fields. The objectives of the proposal are to - combine Argo/CTD profiles and altimetry to calculate temporal and spatial high resolution salinity fields in the mixed layer and in the upper 1500-2000m for the time period 1992 - 2013 for the subpolar and the subtropical Atlantic. - Construct regional freshwater budgets for the mixed layer and the upper 1500-2000m and estimate the contributions of the main processes to the observed change - Follow salinity anomalies from their origin along their circulation pathways, and study involved mechanisms (for instance role of eddies, mean circulation) for shallow, intermediate and deep water masses - Investigate changes in the formation of North Atlantic mode waters in the subpolar and subtropical gyre
Cydia pomonella granulovirus (CpGV, Baculoviridae) is one of the most important agents for the control of codling moth (CM, Cydia pomonella, L.) in both biological and integrated pest management. The rapid emergence of resistance against CpGV-M, which was observed in about 40 European CM field populations from 2003 on, could be traced back to a single, dominant, sex-linked gene. Since then, resistance management has been based on mixtures of new CpGV isolates (CpGV-I12, -S), which are able to overcome this resistance. Recently, resistance even to these novel isolates was observed in CM field populations. This resistance does not follow the described dominant, sex-linked inheritance trait. At the same time, another isolate CpGV-V15 was identified showing high virulence against these resistant populations. To elucidate this novel resistance mechanism and to identify the resistance gene(s) involved, we propose a comprehensive analysis of this resistance on the cellular and genomic level of codling moth. Because of the lack of previous knowledge of the molecular mechanisms of virus resistance in insects, several different and complementary approaches will be pursued. This study will not only give an in-depth insight into the genetic possibilities for development of baculovirus resistance in CM field populations and how the virus overcomes it, but can also serve as an important model for other baculovirus-host interaction systems.
Changes in agroecosystem management (e.g. landscape diversity, management intensity) affect the natural control of pests. The effects of agricultural change on this ecosystem service, however, are not universal and the mechanisms affecting it remain to be understood. As biological control is effectively the product of networks of interactions between pests and their natural enemies, food web analysis provides a versatile tool to address this gap of knowledge. The proposed project will utilize a molecular food web approach and examine, for the first time, how changes in plant fertilisation and landscape complexity affect quantitative aphid-parasitoid-hyperparasitoid food webs on a species-specific level to unravel how changes in food web interactions affect parasitoid aphid control. Based on the fieldderived data, cage experiments will be conducted to assess how parasitoid diversity and identity affect parasitoid interactions and pest control, complementing the field results. The work proposed here will take research on parasitoid aphid control one step further, as it will provide a clearer understanding of how plant fertilization affects whole aphid-parasitoid food webs in both simple and complex landscapes, allowing for further improvements in natural pest control.
The decomposition of terrestrial organic material such as leaf litter represents a fundamental ecosystem function in streams that delivers energy for local and downstream food webs. Although agriculture dominates most regions in Europe and fungicides are applied widely, effects of currently used fungicides on the aquatic decomposer community and consequently the leaf decomposition rate are largely unknown. Also potential compensation of such hypothesised adverse effects due to nutrients or higher average water temperatures associated with climate change are not considered. Moreover, climate change is predicted to alter the community of aquatic decomposers and an open question is, whether this alteration impacts the leaf decomposition rate. The current projects follows a tripartite design to answer these research questions. Firstly, a field study in a vine growing region where fungicides are applied in large amounts will be conducted to whether there is a dose-response relationship between the exposure to fungicides and the leaf decomposition rate. Secondly, experiments in artificial streams with field communities will be carried out to assess potential compensatory mechanisms of nutrients and temperature for effects of fungicides. Thirdly, field experiments with communities exhibiting a gradient of taxa sensitive to climate change will be used to investigate potential climate-related effects on the leaf decomposition rate.
Salinity reduces the productivity of cucumber (Cucumis sativus L.) through osmotic and ionic effects. For given atmospheric conditions we hypothesize the existence of an optimal canopy structure at which water use efficiency is maximal and salt accumulation per unit of dry matter production is minimal. This canopy structure optimum can be predicted by integrating physiological processes over the canopy using a functional-structural plant model (FSPM). This model needs to represent the influence of osmotic stress on plant morphology and stomatal conductance, the accumulation of toxic ions and their dynamics in the different compartments of the system, and their toxic effects in the leaf. Experiments will be conducted to parameterize an extended cucumber FSPM. In in-silico experiments with the FSPM we attempt to identify which canopy structure could lead to maximum long-term water use efficiency with minimum ionic stress. The results from in-silico experiments will be evaluated by comparing different canopy structures in greenhouses. Finally, the FSPM will be used to investigate to which extent the improvement of individual mechanisms of salt tolerance like reduced sensitivity of stomatal conductance or leaf expansion can contribute to whole-plant salt tolerance.
Most soils develop distinct soil architecture during pedogenesis and soil organic carbon (SOC) is sequestered within a hierarchical system of mineral-organic associations and aggregates. Permafrost soils store large amounts of carbon due to their permanently frozen subsoil and a lack of oxygen in the active layer, but they lack complex soil structure. With permafrost thaw more oxidative conditions and increasing soil temperature presumably enhance the build-up of more complex units of soil architecture and may counterbalance, at least partly, SOC mineralization. We aim to explore the development of mineral-organic associations and aggregates under different permafrost impact with respect to SOC stabilization. This information will be linked to environmental control factors relevant for SOC turnover at the pedon and stand scale to bridge processes occurring at the aggregate scale to larger spatial dimensions. We will combine in situ spectroscopic techniques with fractionation approaches and identify mechanisms relevant for SOC turnover at different scales by multivariate statistics and variogram analyses. From this we expect a deeper knowledge about soil architecture formation in the transition of permafrost soils to terrestrial soils and a scale-spanning mechanistic understanding of SOC cycling in permafrost regions.
Die Nukleation von Eispartikeln spielt eine wichtige Rolle bei der Wolken- und Niederschlagsbildung, mit Konsequenten für die atmosphärische Chemie, die Wolkenphysik und das Erdklima. Für eine Quantifizierung und Vorhersage des Einflusses von Wolken in Wettervorhersage- und Klimamodellen muss die Bildung von Eispartikeln daher in einer realistischen Art und Weise beschrieben werden. Einer der wichtigen Bildungsmechanismen ist dabei die heterogene Eisnukleation im Immersionsmodus, bei dem Eis an der Oberfläche eines in einem wässrigen Tröpfchen suspendierten Eiskeims - zum Beispiel eines Mineralstaub- Partikels - gebildet wird. Wir werden im Rahmen dieses Forschungsprojekts zahlreiche Gefrierexperimente im Immersionsmodus durchführen. So werden eine Reihe verschiedener, als Aerosolpartikel in der Atmosphäre vorkommende Materialien auf ihre Eisnukleationseigenschaften hin untersucht werden. Insbesondere sollen hier die Temperatur- und Zeitabhängigkeit der von diesen Materialien ausgelösten Eisnukleation quantifiziert werden. Dabei werden wir spezielles Augenmerk auf die systematische Untersuchung der von porösen Materialien ausgelösten Eisnukleation legen. Es sollen sowohl synthetische Materialien wie beispielsweise mesoporöse Silikate untersucht werden, als auch natürlich vorkommende Materialien wie etwa mikroporöse Zeolithe.
For surface soils, the mechanisms controlling soil organic C turnover have been thoroughly investigated. The database on subsoil C dynamics, however, is scarce, although greater than 50 percent of SOC stocks are stored in deeper soil horizons. The transfer of results obtained from surface soil studies to deeper soil horizons is limited, because soil organic matter (SOM) in deeper soil layers is exposed to contrasting environmental conditions (e.g. more constant temperature and moisture regime, higher CO2 and lower O2 concentrations, increasing N and P limitation to C mineralization with soil depth) and differs in composition compared to SOM of the surface layer, which in turn entails differences in its decomposition. For a quantitative analysis of subsoil SOC dynamics, it is necessary to trace the origins of the soil organic compounds and the pathways of their transformations. Since SOM is composed of various C pools which turn over on different time scales, from hours to millennia, bulk measurements do not reflect the response of specific pools to both transient and long-term change and may significantly underestimate CO2 fluxes. More detailed information can be gained from the fractionation of subsoil SOM into different functional pools in combination with the use of stable and radioactive isotopes. Additionally, soil-respired CO2 isotopic signatures can be used to understand the role of environmental factors on the rate of SOM decomposition and the magnitude and source of CO2 fluxes. The aims of this study are to (i) determine CO2 production and subsoil C mineralization in situ, (ii) investigate the vertical distribution and origin of CO2 in the soil profile using 14CO2 and 13CO2 analyses in the Grinderwald, and to (iii) determine the effect of environmental controls (temperature, oxygen) on subsoil C turnover. We hypothesize that in-situ CO2 production in subsoils is mainly controlled by root distribution and activity and that CO2 produced in deeper soil depth derives to a large part from the mineralization of fresh root derived C inputs. Further, we hypothesize that a large part of the subsoil C is potentially degradable, but is mineralized slower compared with the surface soil due to possible temperature or oxygen limitation.
Glendonites are pseudomorphs after the mineral ikaite (CaCO3 x 6H2O) and composed of calcite (CaCO3). In the past, they have been used as a paleo-thermometer because the primary mineral ikaite, according to observations and experiments, seems to be formed at temperatures near freezing, high alkalinity and high phosphate concentrations in marine sediments. An enigmatic occurrence of the largest glendonites known world-wide, in the Early Eocene Fur Formation of northwestern Denmark offers the unique possibility to shed more light on the actual mechanism and controlling parameters of ikaite formation. Right in the aftermath of the Paleocene-Eocene thermal maximum, a time known for its global pertubation in the global carbon cycle, the formation of authigenic calcium carbonate concretions start in the Fur Formation. In a specific stratigraphic interval inbetween these concretions, the glendonites can be found. We will investigate if termperature changes or changes in geochemical parameters of the Danish Basin caused the sudden formation of ikaite during a time interval that was based on known paleoclimatic reconstructions (semi tropic) not favorable for ikaite formation.
Vorkommen, Häufigkeit, chemische Zusammensetzung und Mischungszustand jener Aerosolpartikel in der Erdatmosphäre, an denen sich durch heterogene Nukleation in unterkühlten Wolken Eis bilden kann (Ice Nucleating Particles = INP), werden experimentell untersucht. Diese Informationen sind wichtig für das Verständnis der Niederschlagsbildung, und finden in parametrisierter Form Eingang in meteorologische Modelle zur Vorhersage des Niederschlages. Das Projekt verwendet hierbei im Wesentlichen physikalische Methoden zur Identifikation und Isolation der Partikel aus der Atmosphäre, und nachfolgend elektronenmikroskopische Methoden zur mineralogischen Analyse einzelner Partikel. Die Identifikation jener wenigen Aerosolpartikel (ca. 1 von 10.000 bis 1 von 100.000), die Eisbildungsfähigkeit besitzen, erfolgt, indem eine Aerosolprobe einer Unterkühlung unter 0°C und Wasserdampfübersättigung ausgesetzt wird, und die an INP entstehenden Eiskristalle fotografiert und gezählt werden. Es werden sowohl Aerosolpartikel aus luftgetragenem Aerosol untersucht (aus dem Eiskeimzähler FINCH) wie auch Partikel, die aus einer Luftprobe auf einem Silizium-Probenträger niedergeschlagen und danach als INP identifiziert wurden (Eiskeimzähler FRIDGE). Eine dritte und vierte Methode (Ice-CVI und ISI) isolieren eisbildungsfähige Partikel, indem aus einer angesaugten Probe von Wolkenluft die Eiskristalle strömungstechnisch von den übrigen Luftbestandteilen getrennt werden. Alle Eiskeimproben werden im Rasterelektronenmikroskop auf Größe, Morphologie, Mischungszustand und chemische Zusammensetzung untersucht und die Ergebnisse der verschiedenen Ansätze verglichen. In Feldexperimenten werden Atmosphärenproben verschiedener geographischer Provenienz (Mitteleuropa, Forschungsstation Jungfraujoch, Wüstenstaub, Vulkanstaub) erhalten. In Laborexperimenten wird mit vorher gesammelt und charakterisierten Modellsubstanzen gearbeitet. Weiterhin wird durch tägliche Messungen der Anzahl-Konzentration und Zusammensetzung von Eiskeimen am Taunus Observatorium nahe Frankfurt über einen längeren Zeitraum untersucht, ob es Saisonalitäten, bevorzugte Quellgebiete (z.B. Wüsten, Industrie, etc.) und biologische Einflussfaktoren (z.B. Pollen, Pflanzenabrieb, Bakterien) für das Vorkommen von Eisnuklei gibt.
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