Das Teilprojekt B1 verfolgt den 'östlichen Verbreitungsweg' der Out-of-Africa 2 Migration von Afrika nach Westeurasien. Geoarchäologische Forschung wurden in Schlüsselgebieten der gesamten Region durchgeführt, um die archäologische und paläoklimatische Entwicklung der letzten ca.50 ka besser zu verstehen. Neben der Fertigstellung laufender Feldforschungen in der nächsten SFB Phase wird der Fokus auf Korrelationen menschlicher Präsenz mit paläogeographischen Determinanten (Niederschlag, Höhe ü NN, etc.) und deren räumlicher Verbreitung liegen. Dies geschieht in Zusammenarbeit mit Teilprojekt D1 durch eine Rekonstruktion der Bedingungen, welche es Homo sapiens ermöglichten, sich in Europa auszubreiten.
Cherry leaf roll virus (CLRV) is a plant pathogen of economic and ecologic importance. It is globally distributed in a wide range of forest, fruit, and ornamental trees and shrubs. In several areas of cherry and walnut production CLRV causes severe losses in yield and quality. With current reference to the rapid dissemination and strong symptom expression in Finnish birches and the Germany-wide distribution of CLRV in birches and elderberry, we continuously investigate and gradually reveal CLRV transmission pathways as by pollen, seeds or water. However, modes and interactions responsible for the wide intergeneric host transmission as well as for the exceptional CLRV epidemic in Fennoscandia still remain unknown. In this project systematic studies shall investigate biological vectors as a causal agent to finally derive control mechanisms and strategies to avoid new epidemics in different hosts and geographic regions. Detailed monitoring of the invertebrate fauna of birch stands/forests and elderberry plantations in Germany and Finland shall reveal potential vectors to subsequently study them in detail by approved virus detection methods and transmission experiments. Molecular analyses of the CLRV coat protein shall prove its role as a viral determinant for a virus/vector interaction. Consequently, this project essentially will contribute important answers on the CLRV epidemiology, and this will be a key element within the first network of research on plant viral pathogens in forest trees.
Der SFB 806 'Unser Weg nach Europa', eingerichtet im Jahre 2009, erforscht die Menschheits- und Umweltgeschichte von der Entstehung der anatomisch modernen Menschen vor 190 000 Jahren in Afrika und ihrer Einwanderung nach Europa. Der SFB 806 setzt geowissenschaftliche, kulturwissenschaftliche und archäologische Methoden ein, um die naturräumlichen und kulturellen Kontexte der Ausbreitungs- und Rückzugsbewegungen prähistorischer Populationen zu rekonstruieren. Ähnlich wie in den ersten beiden Phasen des SFB 806 unterscheiden sich die zu untersuchenden Zeitrahmen (MIS 6 bis MIS 1) je nach Region: Es begann in Ostafrika vor etwa 190.000 Jahren (während MIS 6) mit dem ersten Auftreten von AMH in Äthiopien. Seine Ausbreitung in den Nahen Osten (und andere afrikanische Regionen) erfolgte - basierend auf Ergebnissen von unterschiedlichen Forschungsfeldern außerhalb des SFB 806 - während MIS 5 bis MIS 3 in mehreren 'Wellen', die nicht alle erfolgreich waren, d.h. mit der lang anhaltende Vorherrschaft unserer Vorfahren in diesen neuen Regionen verbunden waren. Vor allem im Nahen Osten wurde AMH zunächst von Neandertalern 'zurückgedrängt', kehrte aber während des MIS 4 bis MIS 3 zurück, um sich dann weiter nach SO-Europa auszubreiten. MIS 5 bis MIS 2 sind jeweils relevante Zeitrahmen für die Arbeiten in unserem westlichen Korridor, während sich die Ausbreitung von AMH in Mitteleuropa während MIS 3/4 bis MIS 1 vollzog. Was waren die Hauptfaktoren, die diese Mobilität ermöglichten oder verhinderten? Bisher haben wir deutliche Hinweise, dass es sich um eine Reihe von Faktoren gehandelt haben muss, die zum einen gemeinsam vorkamen, sich andererseits aber in Raum und Zeit unterschieden: Klima, Umwelt und kultureller Kontext sowie Bevölkerungsdichten sind nennenswerte Faktoren. Derzeit wird diskutiert (Stringer 2011: 221), ob Bevölkerungswachstum einer der Faktoren für die Akkumulation und die Erhaltung von neuen Verhaltensweisen in alten menschlichen Gemeinschaften waren. Diese neuen Verhaltensstrategien sollte Menschen dazu befähigt haben, optimal an neue Herausforderungen in neuen Regionen angepasst zu sein.
In my project I aim at a better understanding of the evolution of malacostracan crustaceans, which includes very different groups such as mantis shrimps, krill and lobsters. Previous studies on Malacostraca, on extant as well as on fossil representatives, focussed on adult morphology.In contrast to such approaches, I will apply a Palaeo-Evo-Devo approach to shed new light on the evolution of Malacostraca. Palaeo-Evo-Devo uses data of different developmental stages of fossil malacostracan crustaceans, such as larval and juvenile stages. With this approach I aim at bridging morphological gaps between the different diverse lineages of modern malacostracans by providing new insights into the character evolution in these lineages.An extensive number of larval and juvenile malacostracans is present in the fossil record, but which have only scarcely been studied. The backbone of this project will be on malacostracans from the Solnhofen Lithographic Limestones (ca. 150 million years old), which are especially well preserved and exhibit minute details. During previous studies, I developed new documentation methods for tiny fossils from these deposits, e.g., fluorescence composite microscopy, and also discovered the first fossil mantis shrimp larvae. For malcostracan groups that do not occur in Solnhofen, I will investigate fossils from other lagerstätten, e.g., Mazon Creek and Bear Gulch (USA), or Montceaules- Mines and La-Voulte-sur-Rhône (France). The main groups in focus are mantis shrimps and certain other shrimps (e.g., mysids, caridoids), as well as the bottom-living ten-footed crustaceans (reptantians). Examples for studied structures are leg details, including the feeding apparatus, but also eyes. The results will contribute to the reconstruction of 3D computer models.The data collected in this project will be used for evaluating the relationships within Malacostraca, but mainly for providing plausible evolutionary scenarios, how the modern malacostracan diversity evolved. With the Palaeo-Evo-Devo approach, I am also able to detect shifts in developmental timing, called heterochrony, which is interpreted as one of the major driving forces of evolution. Finally, the reconstructed evolutionary patterns can be compared between the different lineages for convergencies. These comparisons might help to explain the convergent adaptation to similar ecological niches in different malacostracan groups, e.g., life in the deep sea, life on the sea bottom, evolution of metamorphosis or of predatory larvae.As the project requires the investigation of a large number of specimens in different groups, I will assign distinct sub-projects to three doctoral researchers. The results of this project will not only be published in peer-reviewed journals, but will also be presented to the non-scientific public, e.g., during fossil fairs or museum exhibitions with 3D models engraved in glass blocks.
The aim of this project is to co-estimate models of the core and ionosphere magnetic fields, with the longer-term view of building a 'comprehensive' model of the Earths magnetic field. In this first step we would like to take advantage of the progresses made in the understanding of the ionosphere by global M-I-T modelling to better separate the core and ionospheric signals in satellite data. The magnetic signal generated in the ionosphere is particularly difficult to handle because satellite data provide only information on a very narrow local time window at a time. To get around this difficulty, we would like to apply a technique derived from assimilation methods and that has been already successfully applied in outer-core flow studies. The technique relies on a theoretical model of the ionosphere such as the Upper Atmosphere Model (UAM), where statistics on the deviations from a simple background model are estimated. The derived statistics provided in a covariance matrix format can then be use directly in the magnetic data inversion process to obtain the expected core and ionospheric models. We plan to apply the technique on the German CHAMP satellite data selected for magnetically quiet times. As an output we should obtain a model of the ionospheric magnetic variation field tailored for the selected data and a core-lithosphere field model where possible leakage from ionospheric signals are avoided or at least reduced. The technique can in theory be easily extended to handle the large-scale field generated in the magnetosphere.
Chlorinated ethylenes are prevalent groundwater contaminants. Numerous studies have addressed the mechanism of their reductive dehalogenation during biodegradation and reaction with zero-valent iron. However, despite insight with purified enzymes and well-characterized chemical model systems, conclusive evidence has been missing that the same mechanisms do indeed prevail in real-world transformations. While dual kinetic isotope effect measurements can provide such lines of evidence, until now this approach has not been possible for chlorinated ethylenes because an adequate method for continuous flow compound specific chlorine isotope analysis has been missing. This study attempts to close this prevalent research gap by a combination of two complementary approaches. (1) A novel analytical method to measure isotope effects for carbon and chlorine. (2) A carefully chosen set of well-defined model reactants representing distinct dehalogenation mechanisms believed to be important in real-world systems. Isotope trends observed in biotic and abiotic environmental dehalogenation will be compared to these model reactions, and the respective mechanistic hypotheses will be confirmed or discarded. With this hypothesis-driven approach it is our goal to elucidate for the first timdehalogenation reactions.
Prehistoric pits are filled with ancient topsoil material, which has been preserved there over millennia. A characteristic of these pit fillings is that their colour is different depending on the time the soil material was relocated. Soil colour is the result of soil forming processes and soil properties, and it could therefore indicate the soil characteristics present during that specific period. To the best of our knowledge, no investigation analysed and explained the reasons for these soil colour changes over time. The proposed project will investigate soil parameters from pit fillings of different archaeological periods in the loess area of the Lower Rhine Basin (NW-Germany). It aims to implement the measurement of colour spectra as a novel analytical tool for the rapid analyses of a high number of soil samples: the main goal is to relate highresolution colour data measured by a spectrophotometer to soil parameters that were analysed by conventional pedogenic methods and by mid infrared spectroscopy (MIRS), with a main focus on charred organic matter (BPCAs). This tool would enable us to quantify the variation of soil properties over a timescale of several millennia, during different prehistoric periods at regional scale and for loess soils in general. Detailed information concerning changing soil properties on a regional scale is necessary to determine past soil quality and it helps to increase our understanding of prehistoric soil cultivation practices. Furthermore, these information could also help to increase our understanding about agricultural systems in different archaeological periods.
Atmospheric CO2 concentrations present a repetitive pattern of gradual decline and rapid increase during the last climate cycles, closely related to temperature and sea level change. During the Last Glacial Maximum (LGM; 23-19 kyr BP), when sea level was ca. 120 m below present, the ocean must have stored additionally about 750 Gt carbon. There is consensus that the Southern Ocean represents a key area governing past and present CO2 change. The latter is not only of high scientific but also of socio-economic and political concern since the Southern Ocean provides the potential for an efficient sink of anthropogenic carbon. However, the sensitivity of this carbon sink to climate-change induced reorganizations in wind patterns, ocean circulation, stratification, sea ice extent and biological production remains under debate. Models were not yet able to reproduce the necessary mechanisms involved, potentially due to a lack of the dynamic representation/resolution of atmospheric and oceanic circulation as well as missing carbon cycling. Data on past Southern Ocean hydrography and productivity are mainly from the Atlantic sector, thus do not adequately document conditions in the Pacific sector. This sector is not only the largest part of the Southern Ocean, but it also represents the main drainage area of the marine-based West Antarctic Ice Sheet (WAIS). In the proposed study we aim to generate paleo-data sets with a newly established proxy method from sediment core transects across the Pacific Southern Ocean. This will enhance the baselines for the understanding and modeling of the Southern Ocean's role in carbon cyling, i.e. ocean/atmosphere CO2 exchange and carbon sequestration. It will also allow insight into the response of the WAIS to past warmer than present conditions. Paired isotope measurements (oxygen, silicon) will be made on purified diatoms and radiolarians to describe glacial/interglacial contrasts in physical and nutrient properties at surface and subsurface water depth. This will be used to test (i) the impact of yet unconsidered dust-borne micronutrient deposition on the glacial South Pacific on shifts of primary productivity, Si-uptake rates and carbon export, (ii) the 'silicic-acid leakage' hypothesis (SALH) and (iii) the formation and extent of surface water stratification. Diatom and radiolarian oxygen isotopes will provide information on the timing of surface ocean salinity anomalies resulting from WAIS melt water. Climate model simulations using a complex coupled atmosphere ocean general circulation model (AOGCM) in combination with a sophisticated ocean biogeochemical model including Si-isotopes will be used for comparison with the paleo records. The analysis will cover spatial as well as temporal variability patterns of Southern Ocean hydrography, nutrient cycling and air-sea CO2-exchange. With the help of the climate model we aim to better separate and statistically analyse the individual impacts of ocean circulation and bio
Soil organic matter (SOM) controls large part of the processes occurring at biogeochemical interfaces in soil and may contribute to sequestration of organic chemicals. Our central hypothesis is that sequestration of organic chemicals is driven by physicochemical SOM matrix aging. The underlying processes are the formation and disruption of intermolecular bridges of water molecules (WAMB) and of multivalent cations (CAB) between individual SOM segments or between SOM and minerals in close interaction with hydration and dehydration mechanisms. Understanding the role of these mediated interactions will shed new light on the processes controlling functioning and dynamics of biogeochemical interfaces (BGI). We will assess mobility of SOM structural elements and sorbed organic chemicals via advanced solid state NMR techniques and desorption kinetics and combine these with 1H-NMR-Relaxometry and advanced methods of thermal analysis including DSC, TGADSC- MS and AFM-nanothermal analysis. Via controlled heating/cooling cycles, moistening/drying cycles and targeted modification of SOM, reconstruction of our model hypotheses by computational chemistry (collaboration Gerzabek) and participation at two larger joint experiments within the SPP, we will establish the relation between SOM sequestration potential, SOM structural characteristics, hydration-dehydration mechanisms, biological activity and biogechemical functioning. This will link processes operative on the molecular scale to phenomena on higher scales.
During the first project period we developed a general approach to quantify soil pore structure based on X-ray micro-tomography Vogel et al. (2010) which is applicable at various scales to cover soil pores larger that 0.05 mm in a representative way. Based on this method we generated equivalent network models to numerically simulate flow and transport of dissolved chemicals. The existing network model was extended to handle reactive transport and infiltration processes which are especially critical for matter flux in soil. The results were compared to experimental findings. The original research question 'what does a particle see on its way through soil' could be answered quantitatively for various boundary conditions including steady state flux and infiltration. However, we identified various critical aspects of the proposed modeling concept which will be in the focus of the second period. This includes 1) the spatial arrangement of interfaces having different quality which is crucial for chemical interactions and pore scale water dynamics, 2) the realistic multiphase dynamics at the pore scale which need to reflect the dynamic pressure and movement of trapped non-wetting phase and 3) the parametrization of structural complexity which need to be developed beyond the measurement of continuous Minkowski functions to allow the development of quantitative relations between structure and function. These aspects will be explored in a joint experiments in cooperation with partners within the SPP.
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